Catalog

This catalog lists every species from Persian mythology available for simulation in stdferdowsim — 36 species and 360 demographic models. Each has made-up but population-genetically plausible genomes and demographic histories featuring admixture graphs, ghost populations, extinction-recolonization, and complex reticulate topologies with up to 8 populations.

Species are drawn from Ferdowsi’s Shahnameh (Book of Kings), Zoroastrian cosmology, and broader Persian/Iranian mythology.

Demography schematic diagrams (one per model) are shown below when available. To generate them, run from the repo root: python docs/generate_demography_figures.py (requires demesdraw; install with uv sync --extra docs).

  1. Which chromosome (Genome object)?

  2. Which model of demographic history (DemographicModel object)?

Example: simulate Simurgh samples under the ghost-admixture model:

$ stdferdowsim SimAvi -d GhostAdmixture_3F10 -o simurgh.trees -L 100000 Main:10 Secondary:10

Shahnameh Creatures & Heroes

Simurgh (Simurgh avianus) — The mythical bird of the Alborz mountains, who raised Zal, Rostam’s father. Ancient, wise, a bridge between worlds. In the Shahnameh, the Simurgh is both healer and teacher, connecting otherwise separate lineages — a natural metaphor for admixture.

Genome summary

ID:

SimAvi

Species:

Simurgh avianus

Common name:

Simurgh (mythical bird)

Assembly:

Alborz1.0 (GCA_FERD_SimAvi)

Ploidy:

2

Generation time:

200 years

Population size:

15,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.20e-08

8.00e-08

2

18,000,000

1.20e-08

8.00e-08

3

16,000,000

1.20e-08

8.00e-08

4

14,000,000

1.20e-08

8.00e-08

5

12,000,000

1.20e-08

8.00e-08

6

10,000,000

1.20e-08

8.00e-08

alborz_mitogenome

17,000

3.36e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Simurgh (mythical bird) at constant size N=15000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Simurgh (mythical bird). Modern N=15000, bottleneck 1500 at 2000 gen, ancestral 7500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Simurgh (mythical bird). Main N=15000, Secondary N=9000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Div (Divus mazandarensis) — The demons of Mazandaran, deeply diverged from the Iranian hero lineages. The Div-e Sepid (White Demon) is the most powerful. In stdferdowsim, Divs represent ancient, deeply diverged populations.

Genome summary

ID:

DivMaz

Species:

Divus mazandarensis

Common name:

Div (demon of Mazandaran)

Assembly:

DivSefid1.0 (GCA_FERD_DivMaz)

Ploidy:

2

Generation time:

80 years

Population size:

50,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.00e-08

2.00e-08

2

18,000,000

3.00e-08

2.00e-08

3

16,000,000

3.00e-08

2.00e-08

4

14,000,000

3.00e-08

2.00e-08

5

12,000,000

3.00e-08

2.00e-08

mazand_mitogenome

17,000

8.40e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Div (demon of Mazandaran) at constant size N=50000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Div (demon of Mazandaran). Modern N=50000, bottleneck 5000 at 2000 gen, ancestral 25000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Div (demon of Mazandaran). Main N=50000, Secondary N=30000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Azhdaha (Azhdaha draconius) — The Persian dragon, associated with Mount Damavand. Zahhak, possessed by Ahriman, grew serpents from his shoulders. Long-lived, rare — small Ne, very long generation time.

Genome summary

ID:

AzhDra

Species:

Azhdaha draconius

Common name:

Azhdaha (dragon)

Assembly:

Damavand1.0 (GCA_FERD_AzhDra)

Ploidy:

2

Generation time:

500 years

Population size:

5,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

5.00e-09

3.00e-08

2

18,000,000

5.00e-09

3.00e-08

3

16,000,000

5.00e-09

3.00e-08

4

14,000,000

5.00e-09

3.00e-08

5

12,000,000

5.00e-09

3.00e-08

damavand_mitogenome

17,000

1.40e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Azhdaha (dragon) at constant size N=5000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Azhdaha (dragon). Modern N=5000, bottleneck 500 at 2000 gen, ancestral 2500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Azhdaha (dragon). Main N=5000, Secondary N=3000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Rostam-lineage (Rostam heroicus) — The greatest hero of the Shahnameh, from Zabulestan. His lineage spans many generations: Zal, Rostam, Faramarz. Mixed ancestry (Zal raised by Simurgh, married to Rudabeh of Kabul).

Genome summary

ID:

RosHer

Species:

Rostam heroicus

Common name:

Rostam-lineage (hero dynasty)

Assembly:

Zabulestan1.0 (GCA_FERD_RosHer)

Ploidy:

2

Generation time:

35 years

Population size:

25,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-08

2.50e-08

2

18,000,000

2.00e-08

2.50e-08

3

16,000,000

2.00e-08

2.50e-08

4

14,000,000

2.00e-08

2.50e-08

5

12,000,000

2.00e-08

2.50e-08

zabul_mitogenome

17,000

5.60e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Rostam-lineage (hero dynasty) at constant size N=25000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Rostam-lineage (hero dynasty). Modern N=25000, bottleneck 2500 at 2000 gen, ancestral 12500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Rostam-lineage (hero dynasty). Main N=25000, Secondary N=15000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Rakhsh (Rakhsh equinus) — Rostam’s legendary horse, the only steed strong enough to carry him. Small population, unique lineage.

Genome summary

ID:

RakEqu

Species:

Rakhsh equinus

Common name:

Rakhsh (Rostam’s mythical steed)

Assembly:

Rakhsh1.0 (GCA_FERD_RakEqu)

Ploidy:

2

Generation time:

25 years

Population size:

3,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.20e-08

1.50e-08

2

18,000,000

2.20e-08

1.50e-08

3

16,000,000

2.20e-08

1.50e-08

4

14,000,000

2.20e-08

1.50e-08

5

12,000,000

2.20e-08

1.50e-08

rakhsh_mitogenome

17,000

6.16e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Rakhsh (Rostam’s mythical steed) at constant size N=3000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Rakhsh (Rostam’s mythical steed). Modern N=3000, bottleneck 300 at 2000 gen, ancestral 1500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Rakhsh (Rostam’s mythical steed). Main N=3000, Secondary N=1800. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Zal (Zal albinus) — Born with white hair, abandoned on the Alborz, raised by Simurgh. A living admixture between human and divine lineages.

Genome summary

ID:

ZalAlb

Species:

Zal albinus

Common name:

Zal (white-haired, raised by Simurgh)

Assembly:

ZalNest1.0 (GCA_FERD_ZalAlb)

Ploidy:

2

Generation time:

35 years

Population size:

20,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.80e-08

4.00e-08

2

18,000,000

1.80e-08

4.00e-08

3

16,000,000

1.80e-08

4.00e-08

4

14,000,000

1.80e-08

4.00e-08

5

12,000,000

1.80e-08

4.00e-08

zal_mitogenome

17,000

5.04e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Zal (white-haired, raised by Simurgh) at constant size N=20000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Zal (white-haired, raised by Simurgh). Modern N=20000, bottleneck 2000 at 2000 gen, ancestral 10000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Zal (white-haired, raised by Simurgh). Main N=20000, Secondary N=12000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Rudabeh (Rudabeh simurghica) — Princess of Kabul, descendant of Zahhak, married to Zal. Her lineage carries both Iranian and foreign ancestry.

Genome summary

ID:

RudSim

Species:

Rudabeh simurghica

Common name:

Rudabeh-lineage (Kabul princess dynasty)

Assembly:

Kabul1.0 (GCA_FERD_RudSim)

Ploidy:

2

Generation time:

30 years

Population size:

18,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.20e-08

3.00e-08

2

18,000,000

2.20e-08

3.00e-08

3

16,000,000

2.20e-08

3.00e-08

4

14,000,000

2.20e-08

3.00e-08

5

12,000,000

2.20e-08

3.00e-08

kabul_mitogenome

17,000

6.16e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Rudabeh-lineage (Kabul princess dynasty) at constant size N=18000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Rudabeh-lineage (Kabul princess dynasty). Modern N=18000, bottleneck 1800 at 2000 gen, ancestral 9000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Rudabeh-lineage (Kabul princess dynasty). Main N=18000, Secondary N=10800. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Esfandiyar (Esfandiyar smithius) — The bronze-bodied warrior, invulnerable except for his eyes. Kayanian dynasty.

Genome summary

ID:

EsfSmi

Species:

Esfandiyar smithius

Common name:

Esfandiyar-kind (bronze-bodied warrior)

Assembly:

Rouyin1.0 (GCA_FERD_EsfSmi)

Ploidy:

2

Generation time:

30 years

Population size:

32,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-08

2.50e-08

2

18,000,000

2.00e-08

2.50e-08

3

16,000,000

2.00e-08

2.50e-08

4

14,000,000

2.00e-08

2.50e-08

5

12,000,000

2.00e-08

2.50e-08

rouyin_mitogenome

17,000

5.60e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Esfandiyar-kind (bronze-bodied warrior) at constant size N=32000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Esfandiyar-kind (bronze-bodied warrior). Modern N=32000, bottleneck 3200 at 2000 gen, ancestral 16000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Esfandiyar-kind (bronze-bodied warrior). Main N=32000, Secondary N=19200. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Jamshid (Jamshid pishdadius) — King of the Pishdadian dynasty, the golden age of Iran. Eventually overthrown by Zahhak.

Genome summary

ID:

JamPis

Species:

Jamshid pishdadius

Common name:

Jamshid-kind (Pishdadian golden age)

Assembly:

Persepolis1.0 (GCA_FERD_JamPis)

Ploidy:

2

Generation time:

100 years

Population size:

60,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.50e-08

5.00e-08

2

18,000,000

1.50e-08

5.00e-08

3

16,000,000

1.50e-08

5.00e-08

4

14,000,000

1.50e-08

5.00e-08

5

12,000,000

1.50e-08

5.00e-08

persepolis_mitogenome

17,000

4.20e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Jamshid-kind (Pishdadian golden age) at constant size N=60000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Jamshid-kind (Pishdadian golden age). Modern N=60000, bottleneck 6000 at 2000 gen, ancestral 30000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Jamshid-kind (Pishdadian golden age). Main N=60000, Secondary N=36000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Kayanian (Kayanian dynasticus) — The Kayanian dynasty of heroes (Kay Kavus, Kay Khosrow). The heroic age of the Shahnameh.

Genome summary

ID:

KayDyn

Species:

Kayanian dynasticus

Common name:

Kayanian (heroic dynasty)

Assembly:

Kayanian1.0 (GCA_FERD_KayDyn)

Ploidy:

2

Generation time:

30 years

Population size:

45,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-08

3.00e-08

2

18,000,000

2.00e-08

3.00e-08

3

16,000,000

2.00e-08

3.00e-08

4

14,000,000

2.00e-08

3.00e-08

5

12,000,000

2.00e-08

3.00e-08

kayani_mitogenome

17,000

5.60e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Kayanian (heroic dynasty) at constant size N=45000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Kayanian (heroic dynasty). Modern N=45000, bottleneck 4500 at 2000 gen, ancestral 22500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Kayanian (heroic dynasty). Main N=45000, Secondary N=27000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Bahram (Bahram firesteed) — Associated with fire and victory. Bahram V (Bahram Gur) is the hunter-king of legend.

Genome summary

ID:

BahFir

Species:

Bahram firesteed

Common name:

Bahram-kind (fire-associated)

Assembly:

Bahram1.0 (GCA_FERD_BahFir)

Ploidy:

2

Generation time:

30 years

Population size:

35,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.80e-08

3.20e-08

2

18,000,000

2.80e-08

3.20e-08

3

16,000,000

2.80e-08

3.20e-08

4

14,000,000

2.80e-08

3.20e-08

5

12,000,000

2.80e-08

3.20e-08

bahram_mitogenome

17,000

7.84e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Bahram-kind (fire-associated) at constant size N=35000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Bahram-kind (fire-associated). Modern N=35000, bottleneck 3500 at 2000 gen, ancestral 17500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Bahram-kind (fire-associated). Main N=35000, Secondary N=21000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Pahlavan (Pahlavan divanus) — Hero-monster hybrid lineages. Some pahlavans (champions) carry Div blood through ancient admixture.

Genome summary

ID:

PahDiv

Species:

Pahlavan divanus

Common name:

Pahlavan (hero-monster hybrid lineage)

Assembly:

Pahlavan1.0 (GCA_FERD_PahDiv)

Ploidy:

2

Generation time:

35 years

Population size:

28,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.20e-08

3.00e-08

2

18,000,000

2.20e-08

3.00e-08

3

16,000,000

2.20e-08

3.00e-08

4

14,000,000

2.20e-08

3.00e-08

5

12,000,000

2.20e-08

3.00e-08

pahlavan_mitogenome

17,000

6.16e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Pahlavan (hero-monster hybrid lineage) at constant size N=28000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Pahlavan (hero-monster hybrid lineage). Modern N=28000, bottleneck 2800 at 2000 gen, ancestral 14000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Pahlavan (hero-monster hybrid lineage). Main N=28000, Secondary N=16800. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Sorcerer (Sorcerer wizardus) — The jadugar (sorcerers) of the Shahnameh, practitioners of dark arts, often serving Turanian kings.

Genome summary

ID:

SorWiz

Species:

Sorcerer wizardus

Common name:

Sorcerer (jadugar)

Assembly:

Jadoo1.0 (GCA_FERD_SorWiz)

Ploidy:

2

Generation time:

50 years

Population size:

30,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.50e-08

2.50e-08

2

18,000,000

3.50e-08

2.50e-08

3

16,000,000

3.50e-08

2.50e-08

4

14,000,000

3.50e-08

2.50e-08

5

12,000,000

3.50e-08

2.50e-08

jadoo_mitogenome

17,000

9.80e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Sorcerer (jadugar) at constant size N=30000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Sorcerer (jadugar). Modern N=30000, bottleneck 3000 at 2000 gen, ancestral 15000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Sorcerer (jadugar). Main N=30000, Secondary N=18000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Div-e Sepid (Dough giant) — The White Demon, greatest of the Mazandaran Divs, slain by Rostam in the Seven Labours.

Genome summary

ID:

DouGia

Species:

Dough giant

Common name:

Div-e Sepid (White Demon giant)

Assembly:

DivSepid1.0 (GCA_FERD_DouGia)

Ploidy:

2

Generation time:

120 years

Population size:

4,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

8.00e-09

1.50e-08

2

18,000,000

8.00e-09

1.50e-08

3

16,000,000

8.00e-09

1.50e-08

4

14,000,000

8.00e-09

1.50e-08

5

12,000,000

8.00e-09

1.50e-08

sepid_mitogenome

17,000

2.24e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Div-e Sepid (White Demon giant) at constant size N=4000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Div-e Sepid (White Demon giant). Modern N=4000, bottleneck 400 at 2000 gen, ancestral 2000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Div-e Sepid (White Demon giant). Main N=4000, Secondary N=2400. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Afrasiyab (Afrasiyab horrens) — The Turanian warlord, arch-enemy of Iran, grandson of Tur son of Feridun. His lineage represents Turanian-Iranian conflict and occasional admixture.

Genome summary

ID:

AfrHor

Species:

Afrasiyab horrens

Common name:

Afrasiyab-kind (Turanian warlord lineage)

Assembly:

Afrasiyab1.0 (GCA_FERD_AfrHor)

Ploidy:

2

Generation time:

28 years

Population size:

65,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.80e-08

2.20e-08

2

18,000,000

2.80e-08

2.20e-08

3

16,000,000

2.80e-08

2.20e-08

4

14,000,000

2.80e-08

2.20e-08

5

12,000,000

2.80e-08

2.20e-08

afrasiyab_mitogenome

17,000

7.84e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Afrasiyab-kind (Turanian warlord lineage) at constant size N=65000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Afrasiyab-kind (Turanian warlord lineage). Modern N=65000, bottleneck 6500 at 2000 gen, ancestral 32500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Afrasiyab-kind (Turanian warlord lineage). Main N=65000, Secondary N=39000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Turanian (Turanian nomadicus) — The steppe nomad spirits of Turan, the great enemy civilization in the Shahnameh. Large populations, short generation times, representing the mobile steppe peoples.

Genome summary

ID:

TurNom

Species:

Turanian nomadicus

Common name:

Turanian (steppe nomad spirit)

Assembly:

Turan1.0 (GCA_FERD_TurNom)

Ploidy:

2

Generation time:

22 years

Population size:

120,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.00e-08

2.00e-08

2

18,000,000

3.00e-08

2.00e-08

3

16,000,000

3.00e-08

2.00e-08

4

14,000,000

3.00e-08

2.00e-08

5

12,000,000

3.00e-08

2.00e-08

turan_mitogenome

17,000

8.40e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Turanian (steppe nomad spirit) at constant size N=120000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Turanian (steppe nomad spirit). Modern N=120000, bottleneck 12000 at 2000 gen, ancestral 60000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Turanian (steppe nomad spirit). Main N=120000, Secondary N=72000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Spirits, Jinn & Peris

Jinn (Jinnus persicus) — Elemental spirits of pre-Islamic and Islamic Persian tradition. Made of smokeless fire, they exist in a parallel world (Jinnestan). Very large populations, widespread.

Genome summary

ID:

PriJin

Species:

Jinnus persicus

Common name:

Jinn (elemental spirit)

Assembly:

Jinnestan1.0 (GCA_FERD_PriJin)

Ploidy:

2

Generation time:

30 years

Population size:

200,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

5.00e-08

3.00e-08

2

18,000,000

5.00e-08

3.00e-08

3

16,000,000

5.00e-08

3.00e-08

4

14,000,000

5.00e-08

3.00e-08

5

12,000,000

5.00e-08

3.00e-08

jinn_mitogenome

17,000

1.40e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Jinn (elemental spirit) at constant size N=200000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Jinn (elemental spirit). Modern N=200000, bottleneck 20000 at 2000 gen, ancestral 100000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Jinn (elemental spirit). Main N=200000, Secondary N=120000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Peri (Peri paradisi) — Beautiful fairy-like spirits, intermediate between the divine and mortal realms. In the Shahnameh, Peris sometimes interbreed with human lineages — natural admixture events.

Genome summary

ID:

PerPar

Species:

Peri paradisi

Common name:

Peri (fairy/angel-like spirit)

Assembly:

Peristan1.0 (GCA_FERD_PerPar)

Ploidy:

2

Generation time:

40 years

Population size:

100,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

4.00e-08

3.50e-08

2

18,000,000

4.00e-08

3.50e-08

3

16,000,000

4.00e-08

3.50e-08

4

14,000,000

4.00e-08

3.50e-08

5

12,000,000

4.00e-08

3.50e-08

peri_mitogenome

17,000

1.12e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Peri (fairy/angel-like spirit) at constant size N=100000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Peri (fairy/angel-like spirit). Modern N=100000, bottleneck 10000 at 2000 gen, ancestral 50000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Peri (fairy/angel-like spirit). Main N=100000, Secondary N=60000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Al (Al daemonius) — Nocturnal demons from Iranian folklore, associated with sleep paralysis and childhood illness.

Genome summary

ID:

AlDem

Species:

Al daemonius

Common name:

Al (nocturnal demon)

Assembly:

AlNight1.0 (GCA_FERD_AlDem)

Ploidy:

2

Generation time:

20 years

Population size:

70,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

6.00e-08

2.00e-08

2

18,000,000

6.00e-08

2.00e-08

3

16,000,000

6.00e-08

2.00e-08

4

14,000,000

6.00e-08

2.00e-08

al_mitogenome

17,000

1.68e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Al (nocturnal demon) at constant size N=70000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Al (nocturnal demon). Modern N=70000, bottleneck 7000 at 2000 gen, ancestral 35000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Al (nocturnal demon). Main N=70000, Secondary N=42000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Ghoul (Ghoul underworld) — Desert-dwelling undead (ghul), shape-shifters that prey on travellers. Originated in Persian folklore before spreading to Arabic tradition.

Genome summary

ID:

GhoUnd

Species:

Ghoul underworld

Common name:

Ghoul (ghul, desert undead)

Assembly:

Ghul1.0 (GCA_FERD_GhoUnd)

Ploidy:

2

Generation time:

25 years

Population size:

40,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

4.50e-08

1.80e-08

2

18,000,000

4.50e-08

1.80e-08

3

16,000,000

4.50e-08

1.80e-08

4

14,000,000

4.50e-08

1.80e-08

ghul_mitogenome

17,000

1.26e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Ghoul (ghul, desert undead) at constant size N=40000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Ghoul (ghul, desert undead). Modern N=40000, bottleneck 4000 at 2000 gen, ancestral 20000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Ghoul (ghul, desert undead). Main N=40000, Secondary N=24000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Daeva (Daeva warrior) — False gods and war demons in Zoroastrian cosmology, the opposite of the Yazata. Large populations in the armies of Ahriman.

Genome summary

ID:

DaeWar

Species:

Daeva warrior

Common name:

Daeva (false god / war demon)

Assembly:

Daeva1.0 (GCA_FERD_DaeWar)

Ploidy:

2

Generation time:

50 years

Population size:

90,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.50e-08

2.00e-08

2

18,000,000

3.50e-08

2.00e-08

3

16,000,000

3.50e-08

2.00e-08

4

14,000,000

3.50e-08

2.00e-08

5

12,000,000

3.50e-08

2.00e-08

daeva_mitogenome

17,000

9.80e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Daeva (false god / war demon) at constant size N=90000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Daeva (false god / war demon). Modern N=90000, bottleneck 9000 at 2000 gen, ancestral 45000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Daeva (false god / war demon). Main N=90000, Secondary N=54000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Druj (Druj angra) — Demons of deceit and chaos in Zoroastrian theology, the fundamental force of evil (Angra Mainyu).

Genome summary

ID:

DraAng

Species:

Druj angra

Common name:

Druj (demon of deceit/chaos)

Assembly:

Druj1.0 (GCA_FERD_DraAng)

Ploidy:

2

Generation time:

25 years

Population size:

150,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

5.00e-08

1.50e-08

2

18,000,000

5.00e-08

1.50e-08

3

16,000,000

5.00e-08

1.50e-08

4

14,000,000

5.00e-08

1.50e-08

5

12,000,000

5.00e-08

1.50e-08

druj_mitogenome

17,000

1.40e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Druj (demon of deceit/chaos) at constant size N=150000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Druj (demon of deceit/chaos). Modern N=150000, bottleneck 15000 at 2000 gen, ancestral 75000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Druj (demon of deceit/chaos). Main N=150000, Secondary N=90000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Gul-spirit (Gulistan rosagarden) — Guardians of the rose garden (Gulistan), from Saadi’s literary tradition. Representing the gentler spirits of Persian literary mythology.

Genome summary

ID:

GulRos

Species:

Gulistan rosagarden

Common name:

Gul-spirit (rose garden guardian)

Assembly:

Gulistan1.0 (GCA_FERD_GulRos)

Ploidy:

2

Generation time:

20 years

Population size:

55,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

4.00e-08

3.00e-08

2

18,000,000

4.00e-08

3.00e-08

3

16,000,000

4.00e-08

3.00e-08

4

14,000,000

4.00e-08

3.00e-08

5

12,000,000

4.00e-08

3.00e-08

gulistan_mitogenome

17,000

1.12e-07

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Gul-spirit (rose garden guardian) at constant size N=55000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Gul-spirit (rose garden guardian). Modern N=55000, bottleneck 5500 at 2000 gen, ancestral 27500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Gul-spirit (rose garden guardian). Main N=55000, Secondary N=33000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Mythical Birds & Beasts

Huma (Huma paradisica) — The bird of paradise whose shadow bestows kingship. Never lands on the ground. Rare, with long generation times.

Genome summary

ID:

HumBir

Species:

Huma paradisica

Common name:

Huma (bird of paradise)

Assembly:

HumaWing1.0 (GCA_FERD_HumBir)

Ploidy:

2

Generation time:

150 years

Population size:

8,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.00e-08

6.00e-08

2

18,000,000

1.00e-08

6.00e-08

3

16,000,000

1.00e-08

6.00e-08

4

14,000,000

1.00e-08

6.00e-08

5

12,000,000

1.00e-08

6.00e-08

6

10,000,000

1.00e-08

6.00e-08

huma_mitogenome

17,000

2.80e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Huma (bird of paradise) at constant size N=8000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Huma (bird of paradise). Modern N=8000, bottleneck 800 at 2000 gen, ancestral 4000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Huma (bird of paradise). Main N=8000, Secondary N=4800. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Manticore (Manticore persicus) — The martyaxwar (man-eater), a composite creature with a human face, lion body, and scorpion tail. Originally Persian (from mardykhor), a natural metaphor for admixture between deeply diverged lineages.

Genome summary

ID:

ManTig

Species:

Manticore persicus

Common name:

Manticore (martyaxwar)

Assembly:

Martyaxwar1.0 (GCA_FERD_ManTig)

Ploidy:

2

Generation time:

60 years

Population size:

12,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.50e-08

4.00e-08

2

18,000,000

2.50e-08

4.00e-08

3

16,000,000

2.50e-08

4.00e-08

4

14,000,000

2.50e-08

4.00e-08

5

12,000,000

2.50e-08

4.00e-08

mardykhor_mitogenome

17,000

7.00e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Manticore (martyaxwar) at constant size N=12000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Manticore (martyaxwar). Modern N=12000, bottleneck 1200 at 2000 gen, ancestral 6000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Manticore (martyaxwar). Main N=12000, Secondary N=7200. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Senmurv (Senmurv birddog) — A hybrid creature with the body of a dog and the wings and head of a bird. Featured in Sasanian art.

Genome summary

ID:

SenBir

Species:

Senmurv birddog

Common name:

Senmurv (dog-bird hybrid)

Assembly:

Senmurv1.0 (GCA_FERD_SenBir)

Ploidy:

2

Generation time:

45 years

Population size:

10,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-08

5.00e-08

2

18,000,000

2.00e-08

5.00e-08

3

16,000,000

2.00e-08

5.00e-08

4

14,000,000

2.00e-08

5.00e-08

5

12,000,000

2.00e-08

5.00e-08

senmurv_mitogenome

17,000

5.60e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Senmurv (dog-bird hybrid) at constant size N=10000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Senmurv (dog-bird hybrid). Modern N=10000, bottleneck 1000 at 2000 gen, ancestral 5000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Senmurv (dog-bird hybrid). Main N=10000, Secondary N=6000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Chamrosh (Chamrosh griffinus) — Dog-eagle hybrid, disperser of seeds from the world-tree. Related to the Simurgh but smaller.

Genome summary

ID:

ChaGri

Species:

Chamrosh griffinus

Common name:

Chamrosh (dog-eagle, seed-disperser)

Assembly:

Chamrosh1.0 (GCA_FERD_ChaGri)

Ploidy:

2

Generation time:

40 years

Population size:

7,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.50e-08

5.50e-08

2

18,000,000

1.50e-08

5.50e-08

3

16,000,000

1.50e-08

5.50e-08

4

14,000,000

1.50e-08

5.50e-08

5

12,000,000

1.50e-08

5.50e-08

chamrosh_mitogenome

17,000

4.20e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Chamrosh (dog-eagle, seed-disperser) at constant size N=7000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Chamrosh (dog-eagle, seed-disperser). Modern N=7000, bottleneck 700 at 2000 gen, ancestral 3500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Chamrosh (dog-eagle, seed-disperser). Main N=7000, Secondary N=4200. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Shahbaz (Shahbaz shahinus) — The royal falcon, symbol of Persian kingship and hunting tradition. Short generation time, moderate populations.

Genome summary

ID:

ShaSha

Species:

Shahbaz shahinus

Common name:

Shahbaz (royal falcon)

Assembly:

Shahbaz1.0 (GCA_FERD_ShaSha)

Ploidy:

2

Generation time:

15 years

Population size:

6,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.00e-08

6.00e-08

2

18,000,000

3.00e-08

6.00e-08

3

16,000,000

3.00e-08

6.00e-08

4

14,000,000

3.00e-08

6.00e-08

5

12,000,000

3.00e-08

6.00e-08

6

10,000,000

3.00e-08

6.00e-08

shahbaz_mitogenome

17,000

8.40e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Shahbaz (royal falcon) at constant size N=6000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Shahbaz (royal falcon). Modern N=6000, bottleneck 600 at 2000 gen, ancestral 3000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Shahbaz (royal falcon). Main N=6000, Secondary N=3600. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Gavaevodata (Gavaevodata primus) — The primordial bull of Zoroastrian cosmology, from whose body all animals were created after its death. Extremely long generation time, very small Ne — the deepest ancestral lineage.

Genome summary

ID:

GavPri

Species:

Gavaevodata primus

Common name:

Gavaevodata (primordial bull)

Assembly:

PrimalBull1.0 (GCA_FERD_GavPri)

Ploidy:

2

Generation time:

300 years

Population size:

2,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

3.00e-09

2.00e-08

2

18,000,000

3.00e-09

2.00e-08

3

16,000,000

3.00e-09

2.00e-08

4

14,000,000

3.00e-09

2.00e-08

primal_mitogenome

17,000

8.40e-09

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Gavaevodata (primordial bull) at constant size N=2000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Gavaevodata (primordial bull). Modern N=2000, bottleneck 200 at 2000 gen, ancestral 1000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Gavaevodata (primordial bull). Main N=2000, Secondary N=1200. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Zoroastrian Cosmology

Faravahar (Faravahar glorius) — Guardian spirits (fravashi) that protect each person. The winged disc symbol of Zoroastrianism.

Genome summary

ID:

FarGlo

Species:

Faravahar glorius

Common name:

Faravahar (guardian spirit)

Assembly:

Fravashi1.0 (GCA_FERD_FarGlo)

Ploidy:

2

Generation time:

60 years

Population size:

80,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.00e-08

7.00e-08

2

18,000,000

1.00e-08

7.00e-08

3

16,000,000

1.00e-08

7.00e-08

4

14,000,000

1.00e-08

7.00e-08

5

12,000,000

1.00e-08

7.00e-08

fravashi_mitogenome

17,000

2.80e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Faravahar (guardian spirit) at constant size N=80000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Faravahar (guardian spirit). Modern N=80000, bottleneck 8000 at 2000 gen, ancestral 40000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Faravahar (guardian spirit). Main N=80000, Secondary N=48000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Mithra (Mithra solarius) — The deity of covenant and the sun, predating Zoroaster. Later adopted into Roman religion as Mithras. Very long generation time, small populations.

Genome summary

ID:

MitSol

Species:

Mithra solarius

Common name:

Mithra (solar covenant deity)

Assembly:

Mithra1.0 (GCA_FERD_MitSol)

Ploidy:

2

Generation time:

800 years

Population size:

1,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-09

1.00e-08

2

18,000,000

2.00e-09

1.00e-08

3

16,000,000

2.00e-09

1.00e-08

4

14,000,000

2.00e-09

1.00e-08

mithra_mitogenome

17,000

5.60e-09

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Mithra (solar covenant deity) at constant size N=1000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Mithra (solar covenant deity). Modern N=1000, bottleneck 100 at 2000 gen, ancestral 500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Mithra (solar covenant deity). Main N=1000, Secondary N=600. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Yazata (Yazata guardian) — Divine beings worthy of worship in Zoroastrian theology, including Asha (truth), Vohu Manah (good mind), and others. Small populations, deep time.

Genome summary

ID:

YazGua

Species:

Yazata guardian

Common name:

Yazata (divine being worthy of worship)

Assembly:

Yazata1.0 (GCA_FERD_YazGua)

Ploidy:

2

Generation time:

500 years

Population size:

3,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

5.00e-09

2.00e-08

2

18,000,000

5.00e-09

2.00e-08

3

16,000,000

5.00e-09

2.00e-08

4

14,000,000

5.00e-09

2.00e-08

yazata_mitogenome

17,000

1.40e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Yazata (divine being worthy of worship) at constant size N=3000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Yazata (divine being worthy of worship). Modern N=3000, bottleneck 300 at 2000 gen, ancestral 1500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Yazata (divine being worthy of worship). Main N=3000, Secondary N=1800. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Zoroaster-lineage (Zoroaster primus) — The prophet Zarathustra’s spiritual lineage, keepers of the sacred fire.

Genome summary

ID:

ZorPri

Species:

Zoroaster primus

Common name:

Zoroaster-lineage (prophet-fire lineage)

Assembly:

Zartosht1.0 (GCA_FERD_ZorPri)

Ploidy:

2

Generation time:

100 years

Population size:

10,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.20e-08

4.00e-08

2

18,000,000

1.20e-08

4.00e-08

3

16,000,000

1.20e-08

4.00e-08

4

14,000,000

1.20e-08

4.00e-08

5

12,000,000

1.20e-08

4.00e-08

zartosht_mitogenome

17,000

3.36e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Zoroaster-lineage (prophet-fire lineage) at constant size N=10000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Zoroaster-lineage (prophet-fire lineage). Modern N=10000, bottleneck 1000 at 2000 gen, ancestral 5000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Zoroaster-lineage (prophet-fire lineage). Main N=10000, Secondary N=6000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Khidr (Khidr wisdomus) — The Green One, an immortal sage who appears at moments of crisis. The smallest Ne and longest generation time in the catalog — an almost-singleton immortal lineage.

Genome summary

ID:

KhiWis

Species:

Khidr wisdomus

Common name:

Khidr (the Green One, immortal sage)

Assembly:

Khidr1.0 (GCA_FERD_KhiWis)

Ploidy:

2

Generation time:

1000 years

Population size:

500

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

1.00e-09

5.00e-09

2

18,000,000

1.00e-09

5.00e-09

3

16,000,000

1.00e-09

5.00e-09

4

14,000,000

1.00e-09

5.00e-09

khidr_mitogenome

17,000

2.80e-09

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Khidr (the Green One, immortal sage) at constant size N=500.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Khidr (the Green One, immortal sage). Modern N=500, bottleneck 50 at 2000 gen, ancestral 250 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Khidr (the Green One, immortal sage). Main N=500, Secondary N=300. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Cross-cultural & Regional

Nart-spirit (Nart heronius) — From the Nart sagas, the heroic cycle shared between Caucasian and Iranian peoples. Represents cultural and genetic exchange between the Caucasus and Iran.

Genome summary

ID:

NarHer

Species:

Nart heronius

Common name:

Nart-spirit (Caucasian-Iranian hero cycle)

Assembly:

Nart1.0 (GCA_FERD_NarHer)

Ploidy:

2

Generation time:

28 years

Population size:

22,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.50e-08

2.80e-08

2

18,000,000

2.50e-08

2.80e-08

3

16,000,000

2.50e-08

2.80e-08

4

14,000,000

2.50e-08

2.80e-08

5

12,000,000

2.50e-08

2.80e-08

nart_mitogenome

17,000

7.00e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Nart-spirit (Caucasian-Iranian hero cycle) at constant size N=22000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Nart-spirit (Caucasian-Iranian hero cycle). Modern N=22000, bottleneck 2200 at 2000 gen, ancestral 11000 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Nart-spirit (Caucasian-Iranian hero cycle). Main N=22000, Secondary N=13200. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Marduk-spirit (Marduk mortalis) — A Mesopotamian-Iranian crossover spirit, representing the deep cultural exchange between Mesopotamia and the Iranian plateau.

Genome summary

ID:

MarMor

Species:

Marduk mortalis

Common name:

Marduk-spirit (Mesopotamian-Iranian crossover)

Assembly:

Marduk1.0 (GCA_FERD_MarMor)

Ploidy:

2

Generation time:

45 years

Population size:

75,000

Chromosomes

Name

Length (bp)

Mutation rate

Recombination rate

1

20,000,000

2.00e-08

3.50e-08

2

18,000,000

2.00e-08

3.50e-08

3

16,000,000

2.00e-08

3.50e-08

4

14,000,000

2.00e-08

3.50e-08

5

12,000,000

2.00e-08

3.50e-08

marduk_mitogenome

17,000

5.60e-08

0.00e+00

Demographic models

Constant_1F10

Single population constant-size model

Demography schematic

Populations: Main

Single population of Marduk-spirit (Mesopotamian-Iranian crossover) at constant size N=75000.

ThreeEpoch_1F10

Single population three-epoch model with bottleneck

Demography schematic

Populations: Main

Single pop of Marduk-spirit (Mesopotamian-Iranian crossover). Modern N=75000, bottleneck 7500 at 2000 gen, ancestral 37500 at 50000 gen.

TwoPopSplit_2F10

Two population split with symmetric migration

Demography schematic

Populations: Main, Secondary

Two pops of Marduk-spirit (Mesopotamian-Iranian crossover). Main N=75000, Secondary N=45000. Split 15000 gen ago.

GhostAdmixture_3F10

Two sampled pops + extinct ghost with pulse admixture

Demography schematic

Populations: Main, Secondary, Ghost

Ghost contributes 15%% pulse admixture to Main at 5000 gen. Non-tree-like history.

ThreePopAdmix_3F10

Three populations with bidirectional pulse admixture

Demography schematic

Populations: Main, Secondary, Tertiary

10%% Tertiary->Main at 3000 gen, 5%% Main->Secondary at 6000 gen. Admixture graph.

FourRegion_4F10

Four-population stepping-stone across Iranian plateau

Demography schematic

Populations: West, East, North, South

West, East, North, South populations with stepping-stone migration.

FiveRealm_5F10

Five-population admixture graph (Iran-Turan-Rum-Hind-Chin)

Demography schematic

Populations: Iran, Turan, Rum, Hind, Chin

Five Shahnameh realms with pulse admixture events. Tests 5-pop graph inference.

SixPopExtRecolon_6F10

Six-population model with extinction and recolonization

Demography schematic

Populations: Core, Zagros, Alborz, Kavir, Sistan, Recolonized

Kavir goes extinct at 3000 gen, recolonized from Zagros at 2000 gen.

AsymReversal_2F10

Two-pop with migration direction reversal over time

Demography schematic

Populations: Main, Secondary

Migration reverses direction at 5000 gen ago. Tests temporal asymmetry.

SilkRoad_8F10

Eight-population Silk Road stepping-stone model

Demography schematic

Populations: Ray, Isfahan, Shiraz, Tabriz, Mashad, Merv, Balkh, Samarkand

Eight cities along the Silk Road with stepping-stone migration and admixture.

Generic models

In addition to the species-specific models listed in this catalog, stdferdowsim offers generic demographic models that can be run with any species. These are described in the API.